1. Departments of Zoology and Botany; University of British Columbia; Vancouver British Columbia V6T 1Z4 Canada
2. Biodiversity Research Centre; University of British Columbia; Vancouver British Columbia V6T 1Z4 Canada
3. Department of Zoology; Faculty of Resource Science and Technology; Universiti Malaysia Sarawak; 94300 Kota Samarahan Sarawak Malaysia
4. Department of Biology; University of Konstanz; Konstanz Germany
5. Department of Migration and Immuno-ecology; Max Planck Institute for Ornithology; Radolfzell Germany