1. IRD (Research Institute for Development) Centre de Recherche Halieutique Méditerranéenne et Tropicale Avenue Jean Monnet, BP 171, 34203 Sète Cedex, France
2. University of Washington, School of Aquatic and Fishery Sciences 1122 NE Boat Street (Box 355020), Seattle, WA 98195-5020, USA
3. IMARPE (Peruvian Institute of the Sea) Esquina Gamarra y Gral. Valles s/n, Chucuito, Callao, Lima, Peru