1. Scottish Association for Marine Science, Dunstaffnage Marine Laboratory Oban, Argyll PA37 1QA, Scotland, UK
2. Aquatic Systems Group, Agriculture, Food and Environmental Sciences Division, Queen's University Belfast Newforge Lane, BT9 5PX, Northern Ireland, UK
3. Centre for Environment, Fisheries and Aquaculture Science, Burnham Laboratory Remembrance Avenue, Burnham-on-Crouch, Essex CM0 8HA, England, UK
4. FRS Marine Laboratory PO Box 101, 375 Victoria Road, Aberdeen AB11 9DB, Scotland, UK
5. Department of Agricultural and Rural Development Newforge Lane, BT9 5PX, Northern Ireland, UK
6. Joint Nature Conservation Committee Monkstone House, City Road, Peterborough PE1 1JY, England, UK