1. Laboratory of Tropical and Subtropical Agriculture and Ethnobotany, Faculty of Bioscience Engineering, Ghent University, Coupure links 653 , Ghent, 9000, Belgium
2. College of Agriculture and Environmental Sciences, Haramaya University, Po. Box 09 , Dire Dawa, Ethiopia
3. College of Agriculture, Wolaita Sodo University, Po. Box 138 , Sodo, Ethiopia
4. Faculty of Tropical AgriSciences, Czech University of Life Sciences, Kamycka 129 , Prague 6-Suchdol , Prague, 165 21, Czech Republic