1. Department of Microbiology, College of Resource, Sichuan Agricultural University, Yaan 625000, China
2. Xinjiang Production & Construction Corps, Key Laboratory of Protection and Utilization of Biological Resources in Tarim Basin, Tarim University, Alar 843300, China
3. Biotechnology Center, Rice and Sorghum Research Institute, Sichuan Academy of Agricultural Sciences, Luzhou 646100, China
4. Department of Environmental Sciences, University of Helsinki, Helsinki, Fin-00014, Finland