1. 1Costerton Biofilm Center, Department of Immunology and Microbiology, Faculty of Health and Medical Sciences, University of Copenhagen, Copenhagen, Denmark
2. ‡Present address: Department of Biological Sciences, University of Calgary, Calgary, AB, Canada.
3. 2Department of Microbiology, School of Biology, College of Science, University of Tehran, Tehran, Iran
4. 3Singapore Centre for Environmental Life Sciences Engineering, Nanyang Technological University, Singapore