1. School of Biological Sciences, University of East Anglia, Norwich Research Park, Norwich, Norfolk NR4 7TJ, UK
2. Strathclyde Institute of Pharmacy and Biomedical Sciences, University of Strathclyde, 161 Cathedral Street, Glasgow, G4 0RE, UK
3. Department of Biology, Edge Hill University, Lancashire, L39 4QP, UK
4. Department of Molecular Microbiology, John Innes Centre, Norwich, NR4 7UH, UK