1. School of Oral and Dental Sciences, University of Bristol, Lower Maudlin Street, Bristol BS1 2LY, UK
2. School of Biochemistry, University of Bristol, University Walk, Bristol BS8 1TD, UK
3. Department of Oral Biology, University at Buffalo, 223 Foster Hall, Buffalo, NY 14214, USA
4. Center for Oral Health and Systemic Disease, University of Louisville School of Dentistry, 501 South Preston Street, Louisville, KY 40202, USA