1. Strathclyde Institute of Pharmacy and Biomedical Sciences, University of Strathclyde, Glasgow, UK
2. Present address: Ikerbasque, Basque Foundation for Science, Bilbao, Spain
3. Present address: Department of Conservation of Natural Resources, Neiker, Basque Institute for Agricultural Research and Development, Derio, Spain
4. The James Hutton Institute, Invergowrie, Dundee, DD2 5DA, UK