1. Department of Life Science and Research Center for Life Science, College of Science, Rikkyo University, Toshima-ku Nishi-ikebukuro 3-34-1, Tokyo 171-8501, Japan
2. Genome Research Center, NODAI Research Institute, Tokyo University of Agriculture, Setagaya-ku, Sakuragaoka 1-1-1, Tokyo 156-8502, Japan
3. Department of Bioscience, Tokyo University of Agriculture, Setagaya-ku, Sakuragaoka 1-1-1, Tokyo 156-8502, Japan
4. Microbial Genetics Laboratory, Genetic Strains Research Center, National Institute of Genetics, 1111 Yata, Mishima, Shizuoka 411-8540, Japan
5. Department of Genetics, Graduate University for Advanced Studies, Sokendai, 1111 Yata, Mishima, Shizuoka 411-8540, Japan
6. Microbial Function Laboratory, National Food Research Institute, National Agriculture Research Organization, Tsukuba-shi Kannondai 2-1-12, Ibaraki 305-8642, Japan