1. Department of Medical Laboratory Science, Faculty of Health Sciences, Hokkaido University, Sapporo, Japan
2. Division of Biomedical Imaging Research, Juntendo University Graduate School of Medicine, Tokyo, Japan
3. Division of Bioresources Research Center for Zoonosis Control, Hokkaido University, Hokkaido, Sapporo, Japan
4. Laboratory of Microbiology and Immunology, School of Pharmaceutical Sciences, University of Shizuoka, Shizuoka, Japan