1. Department of Life Science and Research Center for Life Science, College of Science, Rikkyo University, Toshima-ku, Tokyo 171-8501, Japan
2. Microbial Genetics Laboratory, Genetic Strains Research Center, National Institute of Genetics, 1111 Yata, Mishima, Shizuoka 411-8540, Japan
3. Department of Biotechnology, Graduate School of Agriculture and Life Sciences, University of Tokyo, Bunkyo-ku, Tokyo 113-8657, Japan
4. Fukushima Medical University, Hiragaoka 1, Fukushima 960-1295, Japan
5. Department of Bioscience, Tokyo University of Agriculture, Setagaya-ku, Tokyo 155-8502, Japan