1. 1School of Biological Sciences, Royal Holloway, University of London, Egham Hill, Egham, Surrey TW20 0EX, UK
2. 2Departamento de Genética, Universidad de Sevilla, Apartado 1095, E-41080 Sevilla, Spain
3. 4Faculty of Biology, University of Freiburg, Schaenzlestr. 1, 79104 Freiburg, Germany
4. 3Biological and Environmental Sciences and Engineering Division, the BioActives lab, King Abdullah University of Science and Technology (KAUST), Thuwal 23955-6900, Kingdom of Saudi Arabia