1. School of Biological Sciences, Life Science Building, The University of Liverpool, PO Box 147, Liverpool L69 7ZB, UK1
2. Department of Biochemistry, University of Otago, Dunedin, New Zealand4
3. Department of Medical Biochemistry2 and Department of Medical Microbiology3, Medical School, University of Cape Town, Observatory 7925, Cape Town, South Africa
4. Institute of Freshwater Ecology, Windermere Laboratories, The Ferry House, Ambleside, Cumbria LA22 0LP, UK5