1. Institute of Infections and Immunity, University of Nottingham, Queen’s Medical Centre, Nottingham NG7 2UH, UK1
2. School of Pharmaceutical Sciences, University of Nottingham, University Park, Nottingham NG7 2RD, UK2
3. School of Biosciences, University of Nottingham, Sutton Bonington Campus, Loughborough LE12 5RD, UK3
4. Immunology Research, R&D 21, VA Medical Center, Portland, OR 97201, USA4