1. Freshwater Aquaculture Collaborative Innovation Center of Hubei Province, Wuhan, Hubei 430070, PR China
2. Department of Aquatic Animal Medicine, College of Fisheries, Huazhong Agricultural University, Wuhan, Hubei 430070, PR China
3. Division of Science and Technology, Beijing Normal University – Hong Kong Baptist University United International College, Zhuhai, Guangdong 519085, PR China
4. School of Biological Sciences, Lake Superior State University, Sault Ste. Marie, MI 49783, USA
5. Institute of Marine and Environmental Technology, University of Maryland, Baltimore, MD 21202, USA
6. Key Lab of Freshwater Animal Breeding, Ministry of Agriculture, Wuhan, Hubei 430070, PR China