1. State Key Laboratory of Virology, Wuhan Institute of Virology, Chinese Academy of Sciences, Wuhan 430071, PR China
2. University of Chinese Academy of Sciences, Beijing 100049, PR China
3. Department of General Surgery, Wuhan No.1 Hospital, Wuhan 430022, PR China
4. Institute for Infection and Immunity, St George's University of London, London SW17 0RE, UK
5. Section of Infectious Diseases, Faculty of Medicine, Imperial College London, St Mary's Campus, London W2 1PG, UK