1. Division of Environmental Science and Ecological Engineering, Korea University, Anam-Dong 5 Ga, Seoul, South Korea
2. Department of Biotechnology, Yonsei University, Sinchon-Dong, Seoul, South Korea
3. Environmental Biotechnology National Core Research Center, Gyeongsang National University, Jinju, South Korea
4. Department of Microbiology, Cornell University, Ithaca, NY 14853-8101, USA