1. University of Exeter in Cornwall, Department of Biosciences, Tremough Campus, Penryn, Cornwall TR10 9EZ, UK
2. University of Bath, Department of Biology and Biochemistry, Claverton Down, Bath BA2 7AY, UK
3. London School of Hygiene and Tropical Medicine, London WC1E 7HT, UK
4. CNRS-UPR9022, Institut de Biologie Moléculaire et Cellulaire, 15 rue René Descartes, 67084 Strasbourg Cedex, France