1. Department of Zoology, University of Oxford, South Parks Road, Oxford OX1 3PS, UK
2. Asia-Pacific Institute of Tropical Medicine and Infectious Diseases, University of Hawaii at Manoa, 3675 Kilauea Avenue, Honolulu, HI 96816, USA
3. Department of Biology, The Pennsylvania State University, University Park, PA 16802, USA
4. Unité des Virus Emergents, Faculté de Médecine de Marseille, Université de la Méditerranée, 27 Boulevard Jean Moulin, 13385 Marseille Cedex 05, France