1. Division of Integrated Biosciences, The University of Tokyo, 202 Bioscience Bldg, 5-1-5 Kashiwanoha, Chiba 277-8562, Japan
2. Division of Agricultural and Life Sciences, The University of Tokyo, 1-1-1 Yayoi, Bunkyo-ku, Tokyo 113-8657, Japan
3. Department of Agricultural Sciences, School of Agriculture, Ibaraki University, Chuo 3-21-1, Ami-machi, Inashiki-gun, Ibaraki 300-0393, Japan