1. Korea Research Institute of Bioscience and Biotechnology, 52 Oeundong, Yusong, Daejeon 305-333, Republic of Korea
2. Division of Environmental Biotechnology, PMBBRC, Gyeongsang National University, Jinju 660-701, Republic of Korea
3. The Key Laboratory for Microbial Resources of Ministry of Education, Yunnan Institute of Microbiology, Yunnan University, Kunming 650091, PR China