1. Korea Research Institute of Bioscience and Biotechnology, 52 Oeundong, Yusong, Daejeon 305-333, Republic of Korea
2. Division of Applied Life Science, EB-NCRC, PMBBRC, Gyeongsang National University, 660-701, Republic of Korea
3. Yunnan Institute of Microbiology, Yunnan University, Kunming, Yunnan, 650091, People's Republic of China