1. College of Bio-resources and Environmental Science, Jishou University, Jishou, Hunan 416000, PR China
2. Yunnan Institute of Microbiology and Key Laboratory for Conservation and Utilization of Bio-resources, Yunnan University, Kunming, Yunnan 650091, PR China
3. DSMZ – Deutsche Sammlung von Mikroorganismen und Zellkulturen GmbH, Inhoffenstraße 7b, D-38124 Braunschweig, Germany
4. Yunnan Salt & Chemical Industry Co., Ltd, Kunming, Yunnan 650091, PR China