1. Department of Bioscience and Technology, School of Engineering, Hokkaido Tokai University, Minamisawa, Minami-ku, Sapporo 005-8601, Japan
2. Bioscience and Chemistry Division, Hokkaido National Industrial Research Institute, Tsukisamu-Higashi, Toyohira-ku, Sapporo 062-8517, Japan
3. Research Laboratory, Higeta Shoyu Co., Ltd, Choshi, Chiba 288-8680, Japan