1. Korea Collection for Type Cultures, Korea Research Institute of Bioscience and Biotechnology, PO Box 115, Yusong, Taejon 305-600, Republic of Korea
2. Department of Microbiology, College of Natural Sciences and Research Center for Molecular Microbiology, Seoul National University, Seoul 151-742, Republic of Korea
3. Department of Agricultural and Environmental Science, University of Newcastle, Newcastle upon Tyne NE1 7RU, UK
4. Department of Biology, College of Natural Sciences, Sunchon National University, Sunchon 540-742, Republic of Korea
5. Division of Life Sciences, Chungbuk National University, Gaeshin-dong, Cheongju, Chungbuk 361-763, Republic of Korea