1. State Key laboratory Breeding Base for Sustainable Control of Pest and Disease, Key Laboratory of Biotechnology, Zhejiang Academy of Agricultural Sciences, Hangzhou 310021, PR China
2. Hunan Provincial Key Laboratory for Biology and Control of Plant Diseases and Insect Pests, College of Plant Protection, Hunan Agricultural University, Changsha 410128, PR China
3. The James Hutton Institute, Invergowrie, Dundee DD2 5DA, UK
4. MRC-University of Glasgow Centre for Virus Research, Glasgow G61 1QH, UK