1. Tokyo University of Agriculture and Technology, Saiwai, Fuchu, Tokyo 183-8509, Japan
2. Invasive Insect Biocontrol and Behavior Laboratory, USDA Agricultural Research Service (USDA-ARS), Beltsville Agricultural Research Center, 10300 Baltimore Avenue, Beltsville, MD 20705, USA
3. Yaeyama Branch Office, Okinawa Prefectural Plant Protection Center, 1178-6, Chisokobaru, Hirae, Ishigaki, Okinawa 907-0003, Japan