1. Department of Biomedicine, Center of Biomedical Research, Graduate School of Medical Sciences, Kyushu University, 3-1-1 Maidashi, Higashi-ku, Fukuoka 812-8582, Japan
2. Center of Biomedical Research, Research Center for Human Disease Modeling, Graduate School of Medical Sciences, Kyushu University, Fukuoka 812-8582, Japan
3. Department of Laboratory Animal Science, Faculty of Agriculture, Tottori University, Tottori 680-8553, Japan
4. Faculty of Life Sciences, Kyoto Sangyo University, Kyoto 603-8555, Japan
5. Laboratory of Laboratory Animal Science and Medicine, Department of Disease Control, Graduate School of Veterinary Medicine, Hokkaido University, Sapporo 060-0818, Japan
6. Division of Molecular Immunology, Institute for Genetic Medicine, Hokkaido University, Sapporo, 060-0815, Japan