1. 1Nuffield Department of Medicine, University of Oxford, Peter Medawar Building, South Parks, Oxford, OX1 3SY, UK
2. 2Department of Microbiology, Faculty of Science, Mahidol University, Bangkok, 10400, Thailand
3. 3Institute of Integrative Biology, University of Liverpool, Biosciences Building, Crown Street, L69 7ZB Liverpool, UK
4. 4Department of Biosystems, Laboratory of Gene Technology, KU Leuven. Kasteelpark Arenberg 21, Box 2462, 3001 Leuven, Belgium
5. 5Departments of Food Science, and Pathobiology, University of Guelph, 50 Stone Rd E, Guelph, ON, N1G 2W1, Canada