1. 1Hunan Provincial Key Laboratory for Biology and Control of Plant Diseases and Insect Pests, Hunan Agricultural University, Changsha, Hunan 410128, PR China
2. 2Institute of Virology, College of Plant Protection, Hunan Agricultural University, Changsha, Hunan 410128, PR China
3. 3Department of Microbiology, 212 Pearson Hall, Miami University, Oxford, OH 45056, USA
4. 4School of Bioscience and Technology, Weifang Medical University, Weifang, Shandong 261053, PR China