1. Sydney Institute of Marine Science, Chowder Bay Road, Mosman, NSW 2088, Australia
2. Department of Biological Sciences, Macquarie University, NSW 2109, Australia
3. School of Biological Sciences, Flinders University, GPO Box 2100, Adelaide, SA 5001, Australia
4. IFREMER, IHPE UMR 5244, Univ. Perpignan Via Domitia, CNRS, Univ. Montpellier, F-34095 Montpellier, France