1. 1Department of Veterinary Microbiology and Preventive Medicine, College of Veterinary Medicine, Iowa State University, Ames, IA, USA
2. 2Department of Entomology, College of Agriculture and Life Sciences, Iowa State University, Ames, Iowa, USA
3. 3Department of Pathobiological Sciences, School of Veterinary Medicine, University of Wisconsin-Madison, Madison, Wisconsin, USA