1. Wildlife Zoonoses and Vector-borne Diseases Research Group, Animal and Plant Health Agency, Addlestone, Surrey, UK
2. School of Veterinary Medicine, University of Surrey, Guildford, UK
3. Virology and Pathogenesis Group, Microbiology Services Division, Public Health England, Wiltshire, UK
4. Department of Clinical Infection, Microbiology and Immunology, University of Liverpool, Liverpool, UK
5. NIHR Health Protection Research Unit in Emerging and Zoonotic Infections, University of Liverpool, Liverpool, UK