1. Zoonosis Science Center, Department of Medical Sciences, Uppsala University, Uppsala, Sweden
2. Zoonosis Science Center, Department of Medical Biochemistry and Microbiology, Uppsala University, Uppsala, Sweden
3. Present address: Sydney Institute for Infectious Diseases, School of Life and Environmental Sciences and School of Medical Sciences, The University of Sydney, New South Wales, Sydney, Australia
4. Department of Pathology and Wildlife Disease, Swedish Veterinary Institute, Uppsala, Sweden
5. Present address: Institute of Environmental Engineering, ETH, Zürich, Switzerland
6. Department of Medical Sciences, Section of Infectious Diseases, Uppsala University, Uppsala, Sweden