1. Laboratory of Systems Virology, Department of Biosystems Science, Institute for Frontier Life and Medical Sciences, Kyoto University, Kyoto, Japan
2. Graduate School of Biostudies, Kyoto University, Kyoto, Japan
3. Graduate School of Pharmaceutical Sciences, Kyoto University, Kyoto, Japan
4. Micro/Nano Technology Center, Tokai University, Kanagawa, Japan
5. Faculty of Science, Kyoto University, Kyoto, Japan
6. Division of Human Genetics, Department of Integrated Genetics, National Institute of Genetics, Shizuoka, Japan
7. Department of Molecular Life Science, Tokai University School of Medicine, Tokai University, Kanagawa, Japan
8. Department of Animal Science, Faculty of Agriculture, Tokyo University of Agriculture, Kanagawa, Japan
9. CREST, Japan Science and Technology Agency, Saitama, Japan
10. Present address: Division of Systems Virology, Department of Infectious Disease Control, Institute of Medical Science, The University of Tokyo, 4-6-1 Shirokanedai, Minato-ku, 1088639 Tokyo, Japan.