1. Department of Aquatic Animal Medicine, Research Center for Marine Biology, College of Fisheries, Huazhong Agricultural University, Wuhan, Hubei 430070, PR China
2. State Key Laboratory of Marine Resource Utilization in the South China Sea, Key Laboratory of Tropical Biological Resources of the Ministry of Education, College of Marine Science, Hainan University, Haikou, Hainan 570228, PR China