1. 1 Wildlife Zoonoses and Vector-borne Diseases Research Group, Animal and Plant Health Agency (APHA), Addlestone, Surrey, KT15 3NB, UK
2. 2 Institute for Infection and Immunity, St George’s Hospital Medical School, University of London, London, UK
3. 3 Department of Clinical Infection, Microbiology and Immunology, Institute of Infection and Global Health, University of Liverpool, Liverpool, UK
4. 4 School of Life Sciences, University of West Sussex, Falmer, West Sussex, UK