1. School of Biological Sciences, University of East Anglia, Norwich Research Park, Norwich, NR4 7TJ, UK
2. Department of Molecular Microbiology, John Innes Centre, Norwich Research Park, Norwich, NR4 7UH, UK
3. Department of Plant Protection, Institute of Industrial and Fodder Crops, Hellenic Agricultural Organization–DEMETER, Volos, Greece
4. Department of Crop Genetics, John Innes Centre, Norwich Research Park, NR4 7UH, Norwich, UK