1. School of Biosciences, College of Health and Life Sciences, Aston University, Aston Triangle, Birmingham B4 7ET, UK
2. School of Biosciences, University of Nottingham, Sutton Bonington Campus, Sutton Bonington, Loughborough LE12 5RD, UK
3. Institute of Microbiology and Infection, School of Biosciences, University of Birmingham, Birmingham B15 2TT, UK