1. School of Biology and Biomedical Sciences Research Complex, University of St Andrews, North Haugh, St Andrews, Fife, KY16 9ST, UK
2. Strathclyde Institute of Pharmacy and Biomedical Sciences, University of Strathclyde, Glasgow, G4 0RE, UK
3. Cell and Molecular Sciences, James Hutton Institute, Invergowrie, Dundee, DD2 5DA, UK