1. Menzies School of Health Research, Charles Darwin University, Darwin, Northern Territory 0811, Australia
2. Research Institute for the Environment and Livelihoods, Charles Darwin University, Darwin, Northern Territory 0811, Australia
3. Berrimah Veterinary Laboratory, Department of Primary Industry, Fisheries and Mines, Berrimah Farm, Makagon Road, Berrimah, Northern Territory 0828, Australia
4. Royal Darwin Hospital and Northern Territory Medical Program, Darwin, Northern Territory 0811, Australia