1. 1Department of Bacteriology and Immunology Lovisenberggata 8, Norwegian Institute of Public Health/Department of Food Safety and Infection Biology, Norwegian University of Life Sciences (NMBU),Oslo,Norway
2. 2Department of Bacteriology - Aquatic and Terrestrial Animals, Norwegian Veterinary Institute (NVI), Oslo, Norway
3. 3Department of Biology and Biochemistry, University of Bath,Claverton Down, Bath,United Kingdom
4. 4Department of Food Safety and Infection Biology, Norwegian University of Life Sciences (NMBU),Oslo,Norway
5. 5Northwest Fisheries Science Center, National Marine Fisheries Service, NOAA,Seattle, WA,United States