1. The School of Veterinary Medicine and Science, The University of Nottingham, Sutton Bonington Campus, Sutton Bonington, Leicestershire LE12 5RD, UK
2. Nuffield Department of Clinical Laboratory Sciences, Oxford University, John Radcliffe Hospital, Headington, Oxfordshire OX3 9DU, UK
3. Institute for Animal Health, Compton, Berkshire RG20 7NN, UK
4. ARK-Genomics, The Roslin Institute, University of Edinburgh, Easter Bush, Midlothian EH25 9RG, UK