1. Department of Bioscience and Biotechnology, Faculty of Agriculture, Kyushu University, Hakozaki 6-10-1, Fukuoka 812-8581, Japan
2. Department of Biostudies, Kyoto University, Kyoto 6060-8501, Japan
3. Department of Applied Bioresource Science, Faculty of Agriculture, Ehime University, Ehime 790-8566, Japan