1. Department of Pathology and Infectious Diseases, School of Veterinary Medicine, University of Surrey, Guildford, UK
2. German Centre for Infection Research (DZIF), partner site Bonn-Cologne, 38124 Braunschweig, Germany
3. Institute for Medical Microbiology, Immunology and Hygiene, University of Cologne, Cologne, Germany
4. Public Health England, London, UK