1. Department of Microbiology, University of Stellenbosch, Stellenbosch 7600, South Africa
2. Department of Applied Biology and Chemistry, Faculty of Applied Biosciences, Tokyo University of Agriculture, Tokyo 156-8502, Japan
3. Department of Bioscience and Biotechnology, Faculty of Agriculture, Graduate School, Kyushu University, Fukuoka 812-8582, Japan
4. Graduate School of Agriculture and Life Sciences, University of Tokyo, Tokyo 113-8657, Japan