1. Department of Microbial, Biochemical and Food Biotechnology, University of the Free State, Bloemfontein 9300, South Africa
2. Laboratorium of Microbiology, University of Ghent, K. L. Ledeganckstraat 35, Gent B-9000, Belgium
3. Laboratorium of BCCM/LMG Bacteria Collection, University of Ghent, K. L. Ledeganckstraat 35, Gent B-9000, Belgium