1. Division of Integrative Environmental Sciences, Graduate School of Life and Environmental Sciences, University of Tsukuba, Tsukuba, Ibaraki 305-8572, Japan
2. Key Laboratory of Marine Biogenetic Resources, The Third Institute of Oceanography SOA, Xiamen, Fujian, 361005, PR China
3. State Key Laboratory of Ocean Engineering, Shanghai Jiaotong University, Shanghai 200240, PR China
4. College of Life Science & Tech., Shanghai Jiaotong University, Shanghai 200240, PR China