1. Cambridge Infectious Diseases Consortium, University of Cambridge, Department of Veterinary Medicine, Madingley Road, Cambridge CB3 0ES, UK
2. Institute of Zoology, Zoological Society of London, Regent’s Park, London NW1 4RY, UK
3. CSIRO Australian Animal Health Laboratories, Portarlington Road, East Geelong, VIC 3219, Australia
4. Wildlife Division of the Forestry Commission, Accra, Ghana
5. University of Glasgow Centre for Virus Research, Institute of Infection, Immunity and Inflammation, College of Medical, Veterinary and Life Sciences, Garscube Estate, Bearsden Road, Glasgow G61 1QH, UK